Tetrapod tracks in Permo–Triassic eolian beds of southern Brazil (Paraná Basin)

Tetrapod tracks in eolianites are widespread in the fossil record since the late Paleozoic. Among these ichnofaunas, the ichnogenusChelichnusis the most representative of the Permian tetrapod ichnological record of eolian deposits of Europe, North America and South America, where theChelichnusIchnofacies often occurs. In this contribution, we describe five sets of tracks (one of which is preserved in cross-section), representing the first occurrence ofDicynodontipusandChelichnusin the “Pirambóia Formation” of southern Brazil. This unit represents a humid desert in southwestern Pangea and its lower and upper contacts lead us to consider its age as Lopingian–Induan. The five sets of tracks studied were compared with several ichnotaxa and body fossils with appendicular elements preserved, allowing us to attribute these tracks to dicynodonts and other indeterminate therapsids. Even though the “Pirambóia Formation” track record is sparse and sub-optimally preserved, it is an important key to better understand the occupation of arid environments by tetrapods across the Permo–Triassic boundary.