Subordination in mixed‐age bird flocks–a removal study

During the winters of 1979 to 1986, flocks (7–17 per season, 78 total) of Willow Tits Parus montanus (generally consisting of an adult pair and four juveniles) were studied in a subalpine mixed forest in Central Norway. The mean survival rate between December and early April for juveniles (0.32) was significantly lower than that found for adults (0.74). As it has previously been found that adult Willow Tits, being dominant, prevent the juveniles in a flock from using preferred foraging sites, a removal experiment was made to see if the survival of juveniles improved when living in flocks with no adults present. After removal of the adult pair from 13 flocks in early January, the juveniles from these flocks survived significantly less well, up to early April, than either the juveniles in 11 flocks in which the adults were present (controlled for flock size) or those in 14 flocks consisting of an adult pair and four juveniles. Despite a skewed access to the resources in favour of dominant adults, the cost of being a subordinate juvenile is probably outweighed by the improved predator evasion and improved food finding efficiency due to the greater experience of the adult pair possessing the territory.