Tamarack (Pinaceae): previously unrecorded developmental host for pine engraver and southern pine engraver (Coleoptera: Scolytidae)

The pine engraver, Ips pini (Say), is broadly distributed across North America (Lanier 1972; Wood 1982; Seybold et al. 1995) with a host range that includes most species of Pinus L., and in rare cases, species of Picea A. Dietrich (both Pinaceae), within its range (Swaine 1918; Bright 1976; Wood 1982; Furniss and Carolin 1992). Ips pini has been recorded from Pinus banksiana Lamb., Pinus resinosa Ait., and Pinus strobus L. (eastern North America), and from three of four subspecies of Pinus contorta Dougl. ex Loudon [P. c. contorta, P. c. latifolia (Engelm.) Critch., and P. c. murrayana (Balf.) Critch.], Pinus coulteri D. Don, Pinus jeffreyi Balf., Pinus ponderosa Dougl. ex P. and C. Laws., and Pinus flexilis James (western North America) (Furniss and Carolin 1992; Seybold et al. 1995). Hopping (1964) reported I. pini on Picea rubens Sarg., Picea glauca (Moench) Voss, and Picea engelmannii Parry ex Engelm. The adult insect is intermediate in length relative to most Ips spp., ranging from 3.3 to 4.5 mm (Hopping 1964; Bright 1976; Wood 1982; USDA Forest Service 1985).